””’Astyanax””’ is a [[genus]] of [[freshwater]] [[Actinopterygii|ray-finned fishes]] belonging to the [[family (biology)|family]] [[Acestrorhamphidae]], the American characins.<ref name=”Cof family”>{{Cof family|family=Acestrorhamphidae|access-date=9 November 2025}}</ref> Some of these fish, like many of their relatives, are kept as [[aquarium]] pets and known collectively as [[tetra]]s. With around 150 described [[species]] and new ones being described yearly,<ref name=”FB”>{{FishBase_genus | genus = Astyanax | month = October | year = 2016 }}</ref> this genus is among the largest of the entire order; ”[[Hyphessobrycon]]” also has more than 145 species and which one is larger at any one time depends on whether more species have been recently described in one or the other. The blind and colorless cave tetra of Mexico is a famous member of the genus, but its [[Taxonomy (biology)|taxonomic]] position is disputed: Some recognize it as [[Mexican tetra#Blind cave form|part of the Mexican tetra]] (”A. mexicanus”) and this is supported by [[phylogenetic]] evidence,<ref>{{cite journal | author = Jeffery W | year = 2009 | title = Regressive evolution in Astyanax cavefish | journal = Annu. Rev. Genet. | volume = 43 | pages = 25–47 | doi=10.1146/annurev-genet-102108-134216| pmid = 19640230 | pmc = 3594788}}</ref><ref>{{cite journal |vauthors=Dowling TE, Martasian DP, Jeffery WR | year = 2002 | title = Evidence for multiple genetic forms with similar eyeless phenotypes in the blind cavefish, Astyanax mexicanus | journal = Mol. Biol. Evol. | volume = 19 | issue = 4| pages = 446–455 | doi=10.1093/oxfordjournals.molbev.a004100| pmid = 11919286 | doi-access = free }}</ref><ref>{{cite journal|last1=Bradic|first1=Martina|last2=Beerli|first2=Peter|last3=García-de León|first3=Francisco J|last4=Esquivel-Bobadilla|first4=Sarai|last5=Borowsky|first5=Richard L|title=Gene flow and population structure in the Mexican blind cavefish complex (Astyanax mexicanus)|journal=BMC Evolutionary Biology|date=2012|volume=12|issue=1|pages=9|doi=10.1186/1471-2148-12-9|pmid=22269119|pmc=3282648 |doi-access=free }}</ref><ref>{{cite journal |vauthors=Strecker U, Faúndez VH, Wilkens H | year = 2004 | title = Phylogeography of surface and cave Astyanax (Teleostei) from Central and North America based on cytochrome b sequence data | journal = Mol. Phylogenet. Evol. | volume = 33 | issue = 2| pages = 469–481 | doi=10.1016/j.ympev.2004.07.001| pmid = 15336680 }}</ref><ref name=”Keene2016″>{{cite book |author1=Keene |author2=Yoshizawa |author3=McGaugh | title=Biology and Evolution of the Mexican Cavefish | year=2016 | pages=77–87 |publisher=Elsevier Science | isbn=978-0-12-802148-4}}</ref> but others recognize the cave form as a separate species, ”[[Astyanax jordani|A. jordani]]”.<ref name=FB/>
””’Astyanax””’ is a [[genus]] of [[freshwater]] [[Actinopterygii|ray-finned fishes]] belonging to the [[family (biology)|family]] [[Acestrorhamphidae]], the American characins.<ref name=”Cof family”>{{Cof family|family=Acestrorhamphidae|access-date=9 November 2025}}</ref> Some of these fish, like many of their relatives, are kept as [[aquarium]] pets and known collectively as [[tetra]]s. With around 150 described [[species]] and new ones being described yearly,<ref name=”FB”>{{FishBase_genus | genus = Astyanax | month = October | year = 2016 }}</ref> this genus is among the largest of the entire order; ”[[Hyphessobrycon]]” also has more than 145 species and which one is larger at any one time depends on whether more species have been recently described in one or the other. The blind and colorless cave tetra of Mexico is a famous member of the genus, but its [[Taxonomy (biology)|taxonomic]] position is disputed: Some recognize it as [[Mexican tetra#Blind cave form|part of the Mexican tetra]] (”A. mexicanus”) and this is supported by [[phylogenetic]] evidence,<ref>{{cite journal | author = Jeffery W | year = 2009 | title = Regressive evolution in Astyanax cavefish | journal = Annu. Rev. Genet. | volume = 43 | pages = 25–47 | doi=10.1146/annurev-genet-102108-134216| pmid = 19640230 | pmc = 3594788}}</ref><ref>{{cite journal |vauthors=Dowling TE, Martasian DP, Jeffery WR | year = 2002 | title = Evidence for multiple genetic forms with similar eyeless phenotypes in the blind cavefish, Astyanax mexicanus | journal = Mol. Biol. Evol. | volume = 19 | issue = 4| pages = 446–455 | doi=10.1093/oxfordjournals.molbev.a004100| pmid = 11919286 | doi-access = free }}</ref><ref>{{cite journal|last1=Bradic|first1=Martina|last2=Beerli|first2=Peter|last3=García-de León|first3=Francisco J|last4=Esquivel-Bobadilla|first4=Sarai|last5=Borowsky|first5=Richard L|title=Gene flow and population structure in the Mexican blind cavefish complex (Astyanax mexicanus)|journal=BMC Evolutionary Biology|date=2012|volume=12|issue=1|pages=9|doi=10.1186/1471-2148-12-9|pmid=22269119|pmc=3282648 |doi-access=free }}</ref><ref>{{cite journal |vauthors=Strecker U, Faúndez VH, Wilkens H | year = 2004 | title = Phylogeography of surface and cave Astyanax (Teleostei) from Central and North America based on cytochrome b sequence data | journal = Mol. Phylogenet. Evol. | volume = 33 | issue = 2| pages = 469–481 | doi=10.1016/j.ympev.2004.07.001| pmid = 15336680 }}</ref><ref name=”Keene2016″>{{cite book |author1=Keene |author2=Yoshizawa |author3=McGaugh | title=Biology and Evolution of the Mexican Cavefish | year=2016 | pages=77–87 |publisher=Elsevier Science | isbn=978-0-12-802148-4}}</ref> but others recognize the cave form as a separate species, ”[[Astyanax jordani|A. jordani]]”.<ref name=FB/>
The [[type species]] is ”A. argentatus”,<ref name = “Cof family”/> which was formerly regarded as a [[Synonym (taxonomy)|synonym]] of ”A. mexicanus”.<ref name = “Cogf genus”/> The [[genus|generic]] name comes from [[Astyanax]], a character in [[Greek mythology]], who was the son of [[Hector of Troy]]; in homage to this, several [[specific name (zoology)|specific epithet]]s also refer to the ”[[Iliad]]”.<ref name=FB/>
The [[type species]] is ”A. argentatus”,<ref name = “Cof family”/> which was formerly regarded as a [[Synonym (taxonomy)|synonym]] of ”A. mexicanus”.<ref name = “Cogf genus”/> The [[genus|generic]] name comes from [[Astyanax]], a character in [[Greek mythology]], who was the son of [[Hector of Troy]]; to , name ()| =/>
==Taxonomy and systematics==
==Taxonomy and systematics==
Genus of fishes
| Astyanax | |
|---|---|
| Astyanax mexicanus | |
| Scientific classification |
|
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| Order: | Characiformes |
| Family: | Acestrorhamphidae |
| Subfamily: | Acestrorhamphinae |
| Genus: | Astyanax S. F. Baird & Girard, 1854[1] |
| Type species | |
| Astyanax argentatus | |
| Synonyms[1] | |
Astyanax is a genus of freshwater ray-finned fishes belonging to the family Acestrorhamphidae, the American characins.[1] Some of these fish, like many of their relatives, are kept as aquarium pets and known collectively as tetras. With around 150 described species and new ones being described yearly,[2] this genus is among the largest of the entire order; Hyphessobrycon also has more than 145 species and which one is larger at any one time depends on whether more species have been recently described in one or the other. The blind and colorless cave tetra of Mexico is a famous member of the genus, but its taxonomic position is disputed: Some recognize it as part of the Mexican tetra (A. mexicanus) and this is supported by phylogenetic evidence,[3][4][5][6][7] but others recognize the cave form as a separate species, A. jordani.[2]
The type species is A. argentatus,[1] which was formerly regarded as a synonym of A. mexicanus.[8] The generic name comes from Astyanax, a character in Greek mythology, who was the son of Hector of Troy; a reference to the large silver scales of the type species, resembling armor.[9]
Taxonomy and systematics
This genus is more or less defined as in 1917 by Carl H. Eigenmann. The lack of comprehensive phylogenetic analyses dealing with this genus is hindering a thorough review and the relationships remain as indeterminate as the monophyly is doubtful. The results of 2020 study using a combination of molecular and morphological data appeared to confirm that Astyanax is not monophyletic and suggested the creation of two new genera and the resurrection of Psalidodon.[10]
Species
Astyanax contains the following valid species:[11]
- Astyanax abramis (Jenyns, 1842)
- Astyanax acatlanensis Schmitter-Soto, 2017
- Astyanax aeneus (Günther, 1860) (Banded tetra)
- Astyanax altior Hubbs, 1936 (Yucatán tetra)
- Astyanax anai Angulo, Santos, M. López, Langeani & McMahan, 2018
- Astyanax angustifrons (Regan, 1908)
- Astyanax apiaka Ferreira, F. C. T. Lima, Ribeiro, Flausino Junior, Machado & Mirande, 2023
- Astyanax aramburui Protogino, Miquelarena & H. L. López, 2006
- Astyanax argentatus Baird & Girard, 1854 (Texan tetra)
- Astyanax argyrimarginatus Garutti, 1999
- Astyanax bacalarensis Schmitter-Soto, 2017
- Astyanax bagual Bertaco & Vigo, 2015
- Astyanax bahiensis (Steindachner, 1877)
- Astyanax baileyi (Rosen, 1972)
- Astyanax belizianus (Bocourt, 1868) (Belizean tetra)
- Astyanax bimaculatus (Linnaeus, 1758) (Two-spot tetra)
- Astyanax boliviensis Ruiz-C., Román-Valencia, Taphorn, Buckup & Ortega, 2018
- Astyanax bourgeti C. H. Eigenmann 1908
- Astyanax bransfordii (Gill, 1877)
- Astyanax brevimanus (Günther, 1864). (Quiché tetra)
- Astyanax brevirhinus C. H. Eigenmann, 1908
- Astyanax brevoortii (Gill, 1858)
- Astyanax brucutu Zanata, F. C. T. Lima, Di Dario & Gerhard, 2017
- Astyanax caballeroi (Contreras-Balderas & Rivera-Tiellery, 1985)
- Astyanax caroni Ruiz-C, Román-Valencia & Taphorn, 2023
- Astyanax caucanus (Steindachner, 1879)
- Astyanax chaparae Fowler, 1943
- Astyanax clavitaeniatus Garutti, 2003
- Astyanax cocibolca Bussing, 2008
- Astyanax cordovae (Günther, 1880)
- Astyanax courensis Bertaco, Carvalho & Jerep, 2010
- Astyanax cubilhuitz Schmitter-Soto, 2017
- Astyanax cuyuni Ruiz-C, Román-Valencia & Taphorn, 2023
- Astyanax daguae C. H. Eigenmann, 1913
- Astyanax depressirostris Miranda Ribeiro, 1908
- Astyanax dolinae da Graça, C. A. M. Oliveira, F. C. T. Lima, Pains da Silva & Fernandes, 2017
- Astyanax dorioni (Rosen, 1970)
- Astyanax douradilho Bertaco, 2014
- Astyanax elachylepis Bertaco & Lucinda, 2005
- Astyanax embera Ruiz-C., Román-Valencia, Taphorn, Buckup & Ortega, 2018
- Astyanax epiagos Zanata & Camelier, 2008
- Astyanax eremus Ingenito & Duboc, 2014
- Astyanax fasslii (Steindachner, 1915)
- Astyanax filiferus (C. H. Eigenmann, 1913)
- Astyanax finitimus (Bocourt, 1868)
- Astyanax gandhiae Ruiz-C., Román-Valencia, Taphorn, Buckup & Ortega, 2018
- Astyanax garuttii Ruiz-C, Román-Valencia & Taphorn 2023
- Astyanax gisleni Dahl, 1943
- Astyanax goyacensis C. H. Eigenmann, 1908
- Astyanax gracilior C. H. Eigenmann, 1908
- Astyanax guaricana C. A. M. Oliveira, Abilhoa & Pavanelli, 2013
- Astyanax incaicus Tortonese, 1942
- Astyanax integer Myers, 1930
- Astyanax jacobinae Zanata & Camelier, 2008
- Astyanax jenynsii (Steindachner, 1877)
- Astyanax joaovitori Oliveira, Pavanelli & Bertaco, 2017
- Astyanax jordanensis Alcaraz, Pavanelli & Bertaco, 2009
- Astyanax jordani (Hubbs & Innes, 1936)
- Astyanax kennedyi Géry, 1964
- Astyanax keronolepis P. C. Silva, M. C. Malabarba & L. R. Malabarba, 2019
- Astyanax kompi Hildebrand, 1938
- Astyanax kullanderi W. J. E. M. Costa, 1995
- Astyanax lacustris (Lütken, 1875)
- Astyanax leoni Ruiz-C., Román-Valencia & Taphorn, 2023
- Astyanax leopoldi Géry, Planquette & Le Bail, 1988
- Astyanax lineatus (Perugia, 1891)
- Astyanax longior (Cope 1878)
- Astyanax lorien Zanata, Burger & Camelier, 2018
- Astyanax macal Schmitter-Soto, 2017
- Astyanax maculisquamis Garutti & Britski, 1997
- Astyanax maximus (Steindachner, 1876)
- Astyanax megaspilura Fowler, 1944
- Astyanax metae C. H. Eigenmann, 1914
- Astyanax mexicanus (De Filippi, 1853) (Mexican tetra)
- Astyanax microlepis C. H. Eigenmann, 1913
- Astyanax microschemos Bertaco & Lucena, 2006
- Astyanax multidens (Pearson, 1924)
- Astyanax myersi (Fernández-Yépez, 1950)
- Astyanax nasutus Meek, 1907
- Astyanax nicaraguensis C. H. Eigenmann & Ogle, 1907
- Astyanax nobre Dagosta & Marinho, 2022
- Astyanax novae C. H. Eigenmann, 1911
- Astyanax obscurus (Hensel, 1870)
- Astyanax ocotal Valdez-Moreno, Rodiles-Hernández & Schmitter-Soto, 2017
- Astyanax orbignyanus (Valenciennes, 1850)
- Astyanax orstedii (Lütken, 1875)
- Astyanax orthodus C. H. Eigenmann, 1907
- Astyanax panamensis (Günther, 1864) (Panamanian tetra)
- Astyanax pardensis Salgado, 2021
- Astyanax petenensis (Günther, 1864) (Petén tetra)
- Astyanax pirabitira Lucena, Bertaco & Berbigier, 2013
- Astyanax pirapuan Tagliacollo, Britzke, G. S. C. Silva & Benine, 2011
- Astyanax pirauba (Zanata, Birindelli & Moreira, 2010)
- Astyanax poetzschkei Ahl, 1932
- Astyanax procerus Lucena, J. B. Castro & Bertaco, 2013
- Astyanax rioverde Lozano-Vilano & Schmitter-Soto, 2017
- Astyanax rupestris Zanata, Burger & Camelier, 2018
- Astyanax rupununi Fowler, 1914
- Astyanax saltor Travassos, 1960
- Astyanax salvatoris Valdez-Moreno, Lozano-Vilano & Schmitter-Soto, 2017
- Astyanax scintillans Myers, 1928
- Astyanax siapae Garutti, 2003
- Astyanax sincora Burger, Carvalho & Zanata, 2019
- Astyanax stilbe (Cope, 1870)
- Astyanax superbus Myers, 1942
- Astyanax symmetricus C. H. Eigenmann, 1908
- Astyanax tamiahua Schmitter-Soto, 2017
- Astyanax tarpon (C. H. Eigenmann, 1912)
- Astyanax taurorum Lucena, Zaluski & Lucena, 2017
- Astyanax tehuacanensis Schmitter-Soto 2017
- Astyanax totae Ferreira Haluch & Abilhoa, 2005
- Astyanax trierythropterus Manuel Perreira de Godoy, 1970
- Astyanax turmalinensis Triques, Vono & Caiafa, 2003
- Astyanax unitaeniatus Garutti, 1998
- Astyanax utiariti Bertaco & Garutti, 2007
- Astyanax validus Géry, Planquette & Le Bail, 1991
- Astyanax varii Zanata, Burger, Vita & Camelier, 2019
- Astyanax venezuelae Schultz, 1944
- Astyanax villwocki Zarske & Géry, 1999
- Astyanax viridis Salgado, 2021
- Astyanax yariguies (Torres-Mejia, Hernández & Senechal, 2012)
- Synonyms
References
- ^ a b c d e Fricke, Ron; Eschmeyer, William N. & van der Laan, Richard (eds.). “Genera in the family Acestrorhamphidae”. Catalog of Fishes. California Academy of Sciences. Retrieved 9 November 2025.
- ^ a b Froese, Rainer; Pauly, Daniel (eds.). “Species in genus Astyanax“. FishBase. October 2016 version.
- ^ Jeffery W (2009). “Regressive evolution in Astyanax cavefish”. Annu. Rev. Genet. 43: 25–47. doi:10.1146/annurev-genet-102108-134216. PMC 3594788. PMID 19640230.
- ^ Dowling TE, Martasian DP, Jeffery WR (2002). “Evidence for multiple genetic forms with similar eyeless phenotypes in the blind cavefish, Astyanax mexicanus”. Mol. Biol. Evol. 19 (4): 446–455. doi:10.1093/oxfordjournals.molbev.a004100. PMID 11919286.
- ^ Bradic, Martina; Beerli, Peter; García-de León, Francisco J; Esquivel-Bobadilla, Sarai; Borowsky, Richard L (2012). “Gene flow and population structure in the Mexican blind cavefish complex (Astyanax mexicanus)”. BMC Evolutionary Biology. 12 (1): 9. doi:10.1186/1471-2148-12-9. PMC 3282648. PMID 22269119.
- ^ Strecker U, Faúndez VH, Wilkens H (2004). “Phylogeography of surface and cave Astyanax (Teleostei) from Central and North America based on cytochrome b sequence data”. Mol. Phylogenet. Evol. 33 (2): 469–481. doi:10.1016/j.ympev.2004.07.001. PMID 15336680.
- ^ Keene; Yoshizawa; McGaugh (2016). Biology and Evolution of the Mexican Cavefish. Elsevier Science. pp. 77–87. ISBN 978-0-12-802148-4.
- ^ Cite error: The named reference
Cogf genuswas invoked but never defined (see the help page). - ^ Christopher Scharpf (3 October 2025). “Family ACESTRORHAMPHIDAE Eigenmann 1907 (American Tetras)”. The ETYFish Project Fish Name Etymology Database. Retrieved 9 November 2025.
- ^ Guillermo E. Terán; Mauricio F. Benitez; J. Marcos Mirande (2020). “Opening the Trojan horse: phylogeny of Astyanax, two new genera and resurrection of Psalidodon (Teleostei: Characidae)”. Zoological Journal of the Linnean Society. 190 (4): 1217–1234. doi:10.1093/zoolinnean/zlaa019.
- ^ Fricke, Ron; Eschmeyer, William N. & van der Laan, Richard (eds.). “Species in the genus Astyanax“. Catalog of Fishes. California Academy of Sciences. Retrieved 9 November 2025.
