{{short description|Species in mammal family Spalacidae}}
{{Use mdy dates|date=September 2025}}
[[File:Звичайний сліпак Spalax microphthalmus.jpg|thumb|[[Greater blind mole-rat]] (”Spalax microphthalmus”)|alt=Gray blind mole-rat]]
[[Spalacidae]] is a [[family (biology)|family]] of [[mammal]]s in the [[order (biology)|order]] [[Rodentia]] and part of the [[Muroidea]] [[suborder]]. Members of this family are called spalacids, and include [[blind mole-rat]]s, [[bamboo rat]]s, [[Tachyoryctes|mole-rat]]s, and [[zokor]]s. They are found in Asia, eastern Africa, and eastern Europe, primarily in forests, [[shrubland]]s, and [[grassland]]s, though some species can be found in deserts or [[savanna]]s. They range in size from the [[Middle East blind mole-rat]], at {{convert|13|cm|in|0|abbr=on}} with no tail, to the [[large bamboo rat]], at {{convert|48|cm|in|0|abbr=on}} plus a {{convert|20|cm|in|0|abbr=on}} tail. Spalacids are primarily herbivores, and eat roots, [[bulb]]s, [[tuber]]s, grass, and seeds, with some species also eating insects. Few spalacids have population estimates, but four species—the [[giant root-rat]], [[Mehely’s blind mole-rat]], [[Podolsk blind mole-rat]], and [[sandy blind mole-rat]]—are categorized as [[endangered species|endangered]], while the [[Oltenia blind mole-rat]] is categorized as [[critically endangered]].
The 23 extant species of Spalacidae are divided into 7 [[genus|genera]], divided into 4 subfamilies. [[Myospalacinae]] contains 6 species of zokors in 2 genera, [[Rhizomyinae]] contains 4 species of bamboo rats in 2 genera, [[Spalacinae]] contains 11 species of blind mole-rats in 2 genera, and [[Tachyoryctinae]] contains a single genus of 2 mole-rat species. Several extinct prehistoric spalacid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed.<ref name=”PDBSpalacidae”/>
==Spalacids==
==Spalacids==
The following classification is based on the taxonomy described by the reference work ”[[Mammal Species of the World]]” (2005), with augmentation by generally accepted proposals made since using [[molecular phylogenetics|molecular phylogenetic analysis]], as supported by both the IUCN and the [[American Society of Mammalogists]].<ref name=”MSW”/>
===Subfamily Myospalacinae===
===Subfamily Myospalacinae===
{{Main|Myospalacinae}}
{{Main|Myospalacinae}}
|size={{convert|14|–|25|cm|in|0|abbr=on}} long, with no tail<ref name=”AllMam212213″/>
|size={{convert|14|–|25|cm|in|0|abbr=on}} long, with no tail<ref name=”AllMam212213″/>
|habitat=Grassland<ref name=”IUCNAnatolianblindmole-rat”/>
|habitat=Grassland<ref name=”IUCNAnatolianblindmole-rat”/>
|hunting=Roots, tubers, acorns, plant stems, and other plant parts<ref name=”MamWorld1427″/>
|hunting=Roots, , acorns, plant stems, and other plant parts<ref name=”MamWorld1427″/>
|iucn-status=LC |population=Unknown
|iucn-status=LC |population=Unknown
|direction={{population change unknown}}<ref name=”IUCNAnatolianblindmole-rat”/>
|direction={{population change unknown}}<ref name=”IUCNAnatolianblindmole-rat”/>
|image=File:Palestine_Mole-rat_1.jpg |image-size=180px |image-alt=Gray mole-rat
|image=File:Palestine_Mole-rat_1.jpg |image-size=180px |image-alt=Gray mole-rat
|authority-name=[[Alfred Nehring|Nehring]] |authority-year=1898 |authority-not-original=yes
|authority-name=[[Alfred Nehring|Nehring]] |authority-year=1898 |authority-not-original=yes
|range=Middle East and northwestern Africa |range-image= |range-image-size=180px
|range=Middle East and Africa |range-image= |range-image-size=180px
|size={{convert|13|–|22|cm|in|0|abbr=on}} long, with no tail<ref name=”AllMam212213″/>
|size={{convert|13|–|22|cm|in|0|abbr=on}} long, with no tail<ref name=”AllMam212213″/>
|habitat=Shrubland, grassland, and desert<ref name=”IUCNMiddleEastblindmole-rat”/>
|habitat=Shrubland, grassland, and desert<ref name=”IUCNMiddleEastblindmole-rat”/>
|size={{convert|22|–|28|cm|in|0|abbr=on}} long, with no tail<ref name=”AllMam212213″/>
|size={{convert|22|–|28|cm|in|0|abbr=on}} long, with no tail<ref name=”AllMam212213″/>
|habitat=Grassland<ref name=”IUCNBukovinablindmole-rat”/>
|habitat=Grassland<ref name=”IUCNBukovinablindmole-rat”/>
|hunting=Roots, bulbs, tubers, grass, and seeds, as well as insects<ref name=”MamWorld1427″/>
|hunting=Roots, , tubers, grass, and seeds, as well as insects<ref name=”MamWorld1427″/>
|iucn-status=VU |population=Unknown
|iucn-status=VU |population=Unknown
|direction={{population change unknown}}<ref name=”IUCNBukovinablindmole-rat”/>
|direction={{population change unknown}}<ref name=”IUCNBukovinablindmole-rat”/>
===Subfamily Tachyoryctinae===
===Subfamily Tachyoryctinae===
{{Main|Tachyoryctinae}}
{{Species table |no-note=y |genus=[[Tachyoryctes]] |authority-name=[[Eduard Rüppell|Rüppell]] |authority-year=1835 |species-count=two}}
{{Species table |no-note=y |genus=[[Tachyoryctes]] |authority-name=[[Eduard Rüppell|Rüppell]] |authority-year=1835 |species-count=two}}
{{Species table/row
{{Species table/row
Species in mammal family Spalacidae

Spalacidae is a family of mammals in the order Rodentia and part of the Muroidea suborder. Members of this family are called spalacids, and include blind mole-rats, bamboo rats, mole-rats, and zokors. They are found in Asia, eastern Africa, and eastern Europe, primarily in forests, shrublands, and grasslands, though some species can be found in deserts or savannas. They range in size from the Middle East blind mole-rat, at 13 cm (5 in) with no tail, to the large bamboo rat, at 48 cm (19 in) plus a 20 cm (8 in) tail. Spalacids are primarily herbivores, and eat roots, bulbs, tubers, grass, and seeds, with some species also eating insects. Few spalacids have population estimates, but four species—the giant root-rat, Mehely’s blind mole-rat, Podolsk blind mole-rat, and sandy blind mole-rat—are categorized as endangered, while the Oltenia blind mole-rat is categorized as critically endangered.
The 23 extant species of Spalacidae are divided into 7 genera, divided into 4 subfamilies. Myospalacinae contains 6 species of zokors in 2 genera, Rhizomyinae contains 4 species of bamboo rats in 2 genera, Spalacinae contains 11 species of blind mole-rats in 2 genera, and Tachyoryctinae contains a single genus of 2 mole-rat species. Several extinct prehistoric spalacid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed.[1]
Spalacids
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[2]
Subfamily Myospalacinae
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Chinese zokor | E. fontanierii (H. Milne-Edwards, 1867) |
Central China | Size: 16–24 cm (6–9 in) long, plus 3–7 cm (1–3 in) tail[3]
Habitat: Shrubland and grassland[4] Diet: Roots and grains[5] |
LC
Unknown |
| Rothschild’s zokor | E. rothschildi (Thomas, 1911) |
Central China | Size: 14–18 cm (6–7 in) long, plus 2–4 cm (1–2 in) tail[3]
Habitat: Forest, shrubland, and grassland[6] Diet: Roots and grains[5] |
LC
Unknown |
| Smith’s zokor | E. smithii (Thomas, 1911) |
Central China | Size: 16–25 cm (6–10 in) long, plus 3–4 cm (1–2 in) tail[3]
Habitat: Grassland[7] Diet: Roots and grains[5] |
LC
Unknown |
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| False zokor | M. aspalax (Pallas, 1776) |
Mongolia, southern Russia, and northern China | Size: 16–21 cm (6–8 in) long, plus 3–7 cm (1–3 in) tail[3]
Habitat: Forest and shrubland[8] Diet: Roots and grains[5] |
LC
Unknown |
| Siberian zokor | M. myospalax (Laxmann, 1773) |
Kazakhstan, southern Russia, and northern China |
Size: 20–27 cm (8–11 in) long, plus about 5 cm (2 in) tail[3]
Habitat: Grassland[9] Diet: Roots and grains[5] |
LC
Unknown |
| Transbaikal zokor | M. psilurus (H. Milne-Edwards, 1874) |
Mongolia, southern Russia, and northern China | Size: 20–27 cm (8–11 in) long, plus 3–6 cm (1–2 in) tail[3]
Habitat: Shrubland, grassland, and desert[10] Diet: Roots and grains[5] |
LC
Unknown |
Subfamily Rhizomyinae
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Lesser bamboo rat | C. badius (Hodgson, 1841) |
Southeastern Asia |
Size: 14–26 cm (6–10 in) long, plus 4–10 cm (2–4 in) tail[11]
Habitat: Forest[12] Diet: Shrubs, shoots, and roots[13] |
LC
Unknown |
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Chinese bamboo rat | R. sinensis Gray, 1831 |
China, Myanmar, and Vietnam |
Size: 22–45 cm (9–18 in) long, plus 5–10 cm (2–4 in) tail[3]
Habitat: Forest[14] Diet: Bamboo roots, as well as grass, seeds, and fruit[15] |
LC
Unknown |
| Hoary bamboo rat | R. pruinosus Blyth, 1851 |
India and southeastern Asia |
Size: 24–35 cm (9–14 in) long, plus 9–13 cm (4–5 in) tail[3]
Habitat: Forest and grassland[16] Diet: Bamboo roots, as well as grass, seeds, and fruit[15] |
LC
Unknown |
| Large bamboo rat | R. sumatrensis (Raffles, 1821) |
Southeastern Asia |
Size: 26–48 cm (10–19 in) long, plus 10–20 cm (4–8 in) tail[3]
Habitat: Forest[17] Diet: Bamboo roots, as well as grass, seeds, and fruit[15] |
LC
Unknown |
Subfamily Spalacinae
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Anatolian blind mole-rat | N. xanthodon (Nordmann, 1840) |
Western Asia | Size: 14–25 cm (6–10 in) long, with no tail[11]
Habitat: Grassland[18] Diet: Roots, tubers, acorns, plant stems, and other plant parts[19] |
LC
Unknown |
| Lesser blind mole-rat | N. leucodon (Nordmann, 1840) |
Southeastern Europe |
Size: 15–24 cm (6–9 in) long, with no tail[11]
Habitat: Grassland[20] Diet: Roots, tubers, acorns, plant stems, and other plant parts[19] |
LC
Unknown |
| Middle East blind mole-rat | N. ehrenbergi (Nehring, 1898) |
Middle East and northeastern Africa | Size: 13–22 cm (5–9 in) long, with no tail[11]
Habitat: Shrubland, grassland, and desert[21] Diet: Roots, tubers, acorns, plant stems, and other plant parts[19] |
LC
Unknown |
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Bukovina blind mole-rat | S. graecus Nehring, 1898 |
Romania and southwestern Ukraine | Size: 22–28 cm (9–11 in) long, with no tail[11]
Habitat: Grassland[22] Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[19] |
VU
Unknown |
| Giant blind mole-rat | S. giganteus Nehring, 1898 |
Southwestern Russia | Size: 25–35 cm (10–14 in) long, with no tail[11]
Habitat: Shrubland and grassland[23] Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[19] |
LC
Unknown |
| Greater blind mole-rat | S. microphthalmus Güldenstädt, 1770 |
Ukraine and southwestern Russia | Size: 19–31 cm (7–12 in) long, with no tail[11]
Habitat: Shrubland and grassland[24] Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[19] |
LC
Unknown |
| Kazakhstan blind mole-rat | S. uralensis Tiflov & Usov, 1939 |
Kazakhstan | Size: About 31 cm (12 in) long, with no tail[11]
Habitat: Grassland[25] Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[19] |
NT
Unknown |
| Mehely’s blind mole-rat | S. antiquus Méhely, 1909 |
Romania | Size: Unknown[11]
Habitat: Grassland[26] Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[19] |
EN
3,500–3,800 |
| Oltenia blind mole-rat | S. istricus Méhely, 1909 |
Romania | Size: About 24 cm (9 in) long, with no tail[11]
Habitat: Grassland[27] Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[19] |
CR
Unknown |
| Podolsk blind mole-rat | S. zemni Erxleben, 1777 |
Ukraine | Size: 20–31 cm (8–12 in) long, with no tail[11]
Habitat: Forest and grassland[28] Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[19] |
EN
Unknown |
| Sandy blind mole-rat | S. arenarius Reshetnik, 1939 |
Ukraine |
Size: 19–27 cm (7–11 in) long, with no tail[11]
Habitat: Grassland[29] Diet: Roots, bulbs, tubers, grass, and seeds, as well as insects[19] |
EN
Unknown |
Subfamily Tachyoryctinae
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Giant root-rat | T. macrocephalus Rüppell, 1842 |
Ethiopia | Size: 22–31 cm (9–12 in) long, plus 4–7 cm (2–3 in) tail[11]
Habitat: Grassland[30] Diet: Roots, rhizomes, tubers, bulbs, and corms, as well as grass and legumes[31] |
EN
Unknown |
| Northeast African mole-rat | T. splendens (Rüppell, 1836) |
Eastern Africa | Size: 15–27 cm (6–11 in) long, plus 4–10 cm (2–4 in) tail[11]
Habitat: Forest, savanna, shrubland, and grassland[32] Diet: Roots, rhizomes, tubers, bulbs, and corms, as well as grass and legumes[31] |
LC
Unknown |
References
- ^ Cite error: The named reference
PDBSpalacidaewas invoked but never defined (see the help page). - ^ Cite error: The named reference
MSWwas invoked but never defined (see the help page). - ^ a b c d e f g h i Chernasky; Motis; Burgin, pp. 210–211
- ^ a b Smith, A. T.; Johnston, C. H. (2017) [errata version of 2016 assessment]. “Eospalax fontanierii“. IUCN Red List of Threatened Species. 2016: e.T14118A115120816. doi:10.2305/IUCN.UK.2016-3.RLTS.T14118A22277700.en.
- ^ a b c d e f Nowak, p. 1429
- ^ a b Smith, A. T.; Johnston, C. H. (2017) [errata version of 2016 assessment]. “Eospalax rothschildi“. IUCN Red List of Threatened Species. 2016: e.T14121A115121170. doi:10.2305/IUCN.UK.2016-3.RLTS.T14121A22277586.en.
- ^ a b Smith, A. T.; Johnston, C. H. (2017) [errata version of 2016 assessment]. “Eospalax smithii“. IUCN Red List of Threatened Species. 2016: e.T14122A115121321. doi:10.2305/IUCN.UK.2016-3.RLTS.T14122A22277483.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. “Myospalax aspalax“. IUCN Red List of Threatened Species. 2016: e.T14116A115120685. doi:10.2305/IUCN.UK.2016-3.RLTS.T14116A22277071.en.
- ^ a b Cassola, F. (2016). “Myospalax myospalax“. IUCN Red List of Threatened Species. 2016: e.T14119A22277335. doi:10.2305/IUCN.UK.2016-2.RLTS.T14119A22277335.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. “Myospalax psilurus“. IUCN Red List of Threatened Species. 2016: e.T14120A115121026. doi:10.2305/IUCN.UK.2016-3.RLTS.T14120A22277214.en.
- ^ a b c d e f g h i j k l m n Chernasky; Motis; Burgin, pp. 212–213
- ^ a b Aplin, K.; Lunde, D.; Musser, G.; Frost, A.; Molur, S. (2017) [errata version of 2016 assessment]. “Cannomys badius“. IUCN Red List of Threatened Species. 2016: e.T3759A115066803. doi:10.2305/IUCN.UK.2016-3.RLTS.T3759A22274794.en.
- ^ Nowak, p. 1443
- ^ a b Lunde, D.; Aplin, K.; Musser, G. (2017) [errata version of 2016 assessment]. “Rhizomys sinensis“. IUCN Red List of Threatened Species. 2016: e.T19646A115152572. doi:10.2305/IUCN.UK.2016-3.RLTS.T19646A22275131.en.
- ^ a b c Nowak, p. 1442
- ^ a b Aplin, K.; Molur, S. (2017) [errata version of 2016 assessment]. “Rhizomys pruinosus“. IUCN Red List of Threatened Species. 2016: e.T19645A115152385. doi:10.2305/IUCN.UK.2016-3.RLTS.T19645A22274964.en.
- ^ a b Aplin, K.; Lunde, D. (2017) [errata version of 2016 assessment]. “Rhizomys sumatrensis“. IUCN Red List of Threatened Species. 2016: e.T19647A115152803. doi:10.2305/IUCN.UK.2016-3.RLTS.T19647A22275342.en.
- ^ a b Arslan, A.; Gazzard, A.; Matur, F.; Sozen, M. (2023). “Nannospalax xanthodon“. IUCN Red List of Threatened Species. 2023: e.T14327A22276510. doi:10.2305/IUCN.UK.2023-1.RLTS.T14327A22276510.en.
- ^ a b c d e f g h i j k Nowak, p. 1427
- ^ a b Rusin, M. (2024). “Nannospalax leucodon“. IUCN Red List of Threatened Species. 2024: e.T14328A221788646. doi:10.2305/IUCN.UK.2024-2.RLTS.T14328A221788646.en.
- ^ a b Lövy, M.; Gazzard, A. (2023). “Nannospalax ehrenbergi“. IUCN Red List of Threatened Species. 2023: e.T14326A22276839. doi:10.2305/IUCN.UK.2023-1.RLTS.T14326A22276839.en.
- ^ a b Rusin, M. (2024). “Spalax graecus“. IUCN Red List of Threatened Species. 2024: e.T97249856A217345371. doi:10.2305/IUCN.UK.2024-2.RLTS.T97249856A217345371.en.
- ^ a b Kennerley, R.; Formozov, N.; Sheftel, B. (2016). “Spalax giganteus“. IUCN Red List of Threatened Species. 2016: e.T20429A2772339. doi:10.2305/IUCN.UK.2016-2.RLTS.T20429A2772339.en.
- ^ a b Rusin, M. (2024). “Spalax microphthalmus“. IUCN Red List of Threatened Species. 2024: e.T20430A221789991. doi:10.2305/IUCN.UK.2024-2.RLTS.T20430A221789991.en.
- ^ a b Tsytsulina, K. (2017) [errata version of 2016 assessment]. “Spalax uralensis“. IUCN Red List of Threatened Species. 2016: e.T136581A115210023. doi:10.2305/IUCN.UK.2016-3.RLTS.T136581A22276758.en.
- ^ a b Németh, A.; Csorba, G.; Hegyeli, Z. (2024). “Spalax antiquus“. IUCN Red List of Threatened Species. 2024: e.T97250195A221786525. doi:10.2305/IUCN.UK.2024-2.RLTS.T97250195A221786525.en.
- ^ a b Hegyeli, Z.; Csorba, G.; Németh, A. (2024). “Spalax istricus“. IUCN Red List of Threatened Species. 2024: e.T97250154A217345946. doi:10.2305/IUCN.UK.2024-2.RLTS.T97250154A217345946.en.
- ^ a b Rusin, M. (2024). “Spalax zemni“. IUCN Red List of Threatened Species. 2024: e.T42655A91863646. doi:10.2305/IUCN.UK.2024-2.RLTS.T42655A91863646.en.
- ^ a b Rusin, M. (2025) [errata version of 2024 assessment]. “Spalax arenarius“. IUCN Red List of Threatened Species. 2024: e.T20428A270136276. doi:10.2305/IUCN.UK.2024-2.RLTS.T20428A270136276.en.
- ^ a b Lavrenchenko, L.; Kennerley, R. (2017) [errata version of 2016 assessment]. “Tachyoryctes macrocephalus“. IUCN Red List of Threatened Species. 2016: e.T21293A115161321. doi:10.2305/IUCN.UK.2016-3.RLTS.T21293A22276163.en.
- ^ a b Nowak, p. 1445
- ^ a b Cassola, F. (2017). “Tachyoryctes splendens“. IUCN Red List of Threatened Species. 2017: e.T21299A22275532. doi:10.2305/IUCN.UK.2017-2.RLTS.T21299A22275532.en.
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